One day, Usagi Tsukino, clumsy 2nd-year middle school student, stumbles upon a talking cat named Luna. Luna tells her that she is destined to be Sailor Moon, "champion of love and justice", and she must search for the fabled Moon Princess. Usagi finds friends that turn out to be destined senshi as well, and together they fight to save the world from the certain doom brought upon by the Dark Kingdom.
Mamoru is forced to confront Zoisite in a duel over the Rainbow Crystals, but Usagi inadvertently becomes involved.
Usagi and Mamoru remember their former lives as Princess Serenity and Prince Endymion. The Dark Kingdom kidnaps the injured Mamoru, but Queen Beryl executes Zoisite for his attempt to kill him. Luna and Artemis tell the Sailor Guardians about their tragic past.
Minako tries to cheer Usagi up by taking her to a hair salon, leading to the discovery that Tuxedo Mask is still alive, but now very different.
Determined to become more like the princess she is supposed to be, Usagi decides to enlist in Countess Rose's Princess Seminar.
Usagi and her friends visit the Moonlight Course skiing resort, unaware that Kunzite has sabotaged the Moon Princess Contest in order to unmask Sailor Moon.
After discovering that the Moon Princess was once known for her graceful ice skating, Kunzite tries to use this information to trap Sailor Moon. The best skater among the girls, however, turns out to be Makoto. Tuxedo Mask appears to help them, but only because he does not agree with Kunzite's method of involving bystanders.
