One day, Usagi Tsukino, clumsy 2nd-year middle school student, stumbles upon a talking cat named Luna. Luna tells her that she is destined to be Sailor Moon, "champion of love and justice", and she must search for the fabled Moon Princess. Usagi finds friends that turn out to be destined senshi as well, and together they fight to save the world from the certain doom brought upon by the Dark Kingdom.
Luna discovers that she has an admirer, in the form of an overweight tomcat named Rhett Butler. Meanwhile, the Sailor Guardians and Zoisite compete for the last Rainbow Crystal.
Umino misinterprets Usagi's advice on seducing Naru, and begins to adopt the persona of Tuxedo Mask.
Zoisite disguises himself as Sailor Moon to lure Tuxedo Mask and the Sailor Guardians out into the open. The Dark Kingdom injures Tuxedo Mask's identity, but they are foiled by the appearance of Sailor Venus.
Mamoru is forced to confront Zoisite in a duel over the Rainbow Crystals, but Usagi inadvertently becomes involved.
Usagi and Mamoru remember their former lives as Princess Serenity and Prince Endymion. The Dark Kingdom kidnaps the injured Mamoru, but Queen Beryl executes Zoisite for his attempt to kill him. Luna and Artemis tell the Sailor Guardians about their tragic past.
Minako tries to cheer Usagi up by taking her to a hair salon, leading to the discovery that Tuxedo Mask is still alive, but now very different.
